| BlackRock, Inc. | 1,449,671 | $21.2m | 0.0% |
| Caligan Partners LP | 1,435,130 | $21.0m | 1.0% |
| MORGAN STANLEY | 614,320 | $9.0m | 0.0% |
| ACADIAN ASSET MANAGEMENT LLC | 606,544 | $8.9m | 0.0% |
| VANGUARD CAPITAL MANAGEMENT LLC | 572,113 | $8.4m | 0.0% |
| AQR CAPITAL MANAGEMENT LLC | 559,846 | $8.2m | 0.0% |
| BOOTHBAY FUND MANAGEMENT, LLC | 529,640 | $7.7m | 0.1% |
| RENAISSANCE TECHNOLOGIES LLC | 481,417 | $7.0m | 0.0% |
| DIMENSIONAL FUND ADVISORS LP | 451,095 | $6.6m | 0.0% |
| CAPITAL MANAGEMENT CORP /VA | 448,474 | $6.6m | 1.0% |
| DEUTSCHE BANK AG\ | 353,552 | $5.2m | 0.0% |
| GEODE CAPITAL MANAGEMENT, LLC | 331,577 | $4.9m | 0.0% |
| STATE STREET CORP | 311,896 | $4.6m | 0.0% |
| TWO SIGMA INVESTMENTS, LP | 252,964 | $3.7m | 0.0% |
| D. E. Shaw & Co., Inc. | 245,066 | $3.6m | 0.0% |
| GOLDMAN SACHS GROUP INC | 237,722 | $3.5m | 0.0% |
| JACOBS LEVY EQUITY MANAGEMENT, INC | 217,530 | $3.2m | 0.0% |
| Nuveen, LLC | 156,538 | $2.3m | 0.0% |
| Quantinno Capital Management LP | 155,257 | $2.3m | 0.0% |
| UBS Group AG | 145,816 | $2.1m | 0.0% |
| Man Group plc | 140,712 | $2.1m | 0.0% |
| Hillsdale Investment Management Inc. | 138,821 | $2.0m | 0.0% |
| Assenagon Asset Management S.A. | 126,032 | $1.8m | 0.0% |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 113,374 | $1.7m | 0.0% |
| ARROWSTREET CAPITAL, LIMITED PARTNERSHIP | 106,644 | $1.6m | 0.0% |