| FMR LLC | 10,990,535 | $113.2m | 0.0% |
| RTW INVESTMENTS, LP | 6,422,018 | $66.1m | 0.6% |
| BlackRock, Inc. | 4,969,040 | $51.2m | 0.0% |
| STATE STREET CORP | 3,549,823 | $36.6m | 0.0% |
| ACORN CAPITAL ADVISORS, LLC | 3,497,948 | $36.0m | 9.8% |
| Portolan Capital Management, LLC | 3,490,398 | $36.0m | 1.3% |
| Saturn V Capital Management LP | 3,303,686 | $34.0m | 4.9% |
| VANGUARD CAPITAL MANAGEMENT LLC | 2,694,040 | $27.7m | 0.0% |
| GEODE CAPITAL MANAGEMENT, LLC | 1,571,021 | $16.2m | 0.0% |
| MILLENNIUM MANAGEMENT LLC | 1,093,905 | $11.3m | 0.0% |
| Halter Ferguson Financial Inc. | 772,461 | $8.0m | 1.3% |
| NORTHERN TRUST CORP | 492,465 | $5.1m | 0.0% |
| KENNEDY CAPITAL MANAGEMENT LLC | 488,291 | $5.0m | 0.1% |
| VANGUARD PORTFOLIO MANAGEMENT LLC | 437,150 | $4.5m | 0.0% |
| VANGUARD FIDUCIARY TRUST CO | 412,109 | $4.2m | 0.0% |
| Sands Capital Alternatives, LLC | 405,837 | $4.2m | 0.7% |
| GOLDMAN SACHS GROUP INC | 305,216 | $3.1m | 0.0% |
| MARSHALL WACE, LLP | 300,641 | $3.1m | 0.0% |
| BANK OF AMERICA CORP /DE/ | 278,944 | $2.9m | 0.0% |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 227,939 | $2.3m | 0.0% |
| WELLINGTON MANAGEMENT GROUP LLP | 205,745 | $2.1m | 0.0% |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 199,454 | $2.1m | 0.0% |
| Tanager Wealth Management LLP | 198,175 | $2.0m | 0.2% |
| Bank of New York Mellon Corp | 191,701 | $2.0m | 0.0% |
| UBS Group AG | 169,271 | $1.7m | 0.0% |