| Aberdeen Group plc | 2,230,178 | $17.5m | 0.0% |
| Bleichroeder LP | 2,018,190 | $15.9m | 2.8% |
| Octagon Capital Advisors LP | 1,350,000 | $10.6m | 1.0% |
| BOOTHBAY FUND MANAGEMENT, LLC | 1,346,004 | $10.6m | 0.1% |
| BlackRock, Inc. | 1,172,873 | $9.2m | 0.0% |
| VANGUARD CAPITAL MANAGEMENT LLC | 1,116,241 | $8.8m | 0.0% |
| Lynx1 Capital Management LP | 950,466 | $7.5m | 0.9% |
| Driehaus Capital Management LLC | 946,707 | $7.4m | 0.0% |
| Empery Asset Management, LP | 776,093 | $6.1m | 7.1% |
| Affinity Asset Advisors, LLC | 646,576 | $5.1m | 0.2% |
| GEODE CAPITAL MANAGEMENT, LLC | 620,981 | $4.9m | 0.0% |
| MACKENZIE FINANCIAL CORP | 480,997 | $3.8m | 0.0% |
| STATE STREET CORP | 427,325 | $3.4m | 0.0% |
| ACADIAN ASSET MANAGEMENT LLC | 305,759 | $2.4m | 0.0% |
| ArrowMark Colorado Holdings LLC | 274,140 | $2.2m | 0.1% |
| Corient Private Wealth LP | 245,058 | $1.9m | 0.0% |
| TWO SIGMA INVESTMENTS, LP | 237,863 | $1.9m | 0.0% |
| NORTHERN TRUST CORP | 190,188 | $1.5m | 0.0% |
| JPMORGAN CHASE & CO | 180,452 | $1.4m | 0.0% |
| VANGUARD FIDUCIARY TRUST CO | 166,538 | $1.3m | 0.0% |
| VANGUARD PORTFOLIO MANAGEMENT LLC | 161,782 | $1.3m | 0.0% |
| GOLDMAN SACHS GROUP INC | 139,828 | $1.1m | 0.0% |
| Squarepoint Ops LLC | 137,106 | $1.1m | 0.0% |
| MORGAN STANLEY | 130,536 | $1.0m | 0.0% |
| COMMONWEALTH EQUITY SERVICES, LLC | 115,929 | $911,200 | 0.0% |