| Nantahala Capital Management, LLC | 2,802,894 | $13.1m | 0.2% |
| BlackRock, Inc. | 1,413,381 | $6.6m | 0.0% |
| VANGUARD CAPITAL MANAGEMENT LLC | 1,071,481 | $5.0m | 0.0% |
| Octagon Capital Advisors LP | 1,000,000 | $4.7m | 0.4% |
| GEODE CAPITAL MANAGEMENT, LLC | 564,019 | $2.6m | 0.0% |
| Vivo Capital, LLC | 550,000 | $2.6m | 0.2% |
| MARSHALL WACE, LLP | 547,862 | $2.6m | 0.0% |
| Logos Global Management LP | 473,563 | $2.2m | 0.1% |
| STATE STREET CORP | 450,162 | $2.1m | 0.0% |
| Diametric Capital, LP | 374,132 | $1.8m | 0.2% |
| J. Goldman & Co LP | 327,345 | $1.5m | 0.0% |
| DIADEMA PARTNERS LP | 303,676 | $1.4m | 0.2% |
| MILLENNIUM MANAGEMENT LLC | 271,456 | $1.3m | 0.0% |
| Dauntless Investment Group, LLC | 214,000 | $1.0m | 0.8% |
| CITIGROUP INC | 182,654 | $856,647 | 0.0% |
| NORTHERN TRUST CORP | 181,930 | $853,252 | 0.0% |
| VANGUARD PORTFOLIO MANAGEMENT LLC | 169,196 | $793,529 | 0.0% |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 160,039 | $750,583 | 0.0% |
| Jump Financial, LLC | 153,000 | $717,570 | 0.0% |
| VANGUARD FIDUCIARY TRUST CO | 148,965 | $698,646 | 0.0% |
| GOLDMAN SACHS GROUP INC | 145,304 | $681,475 | 0.0% |
| DIMENSIONAL FUND ADVISORS LP | 143,052 | $678,018 | 0.0% |
| Seven Fleet Capital Management LP | 103,400 | $521,136 | 0.1% |
| Caption Management, LLC | 110,000 | $515,900 | 0.0% |
| MORGAN STANLEY | 109,645 | $514,235 | 0.0% |